Two years after the events in The Bourne Identity, Bourne and his girlfriend, Marie Kreutz, are living in Goa, India. Bourne is beginning to recover some of his memories, and he is troubled by disjointed flashbacks of an assassination he carried out in a Berlin hotel. Meanwhile, in Berlin, a CIA officer under Deputy Director Pamela Landy is trading $3 million for the "Neski Files", documents about the theft of $20 million from the CIA seven years earlier. During the exchange, a Russian assassin named Kirill arrives to intercept the selling. He plants two bombs in the basement electrical circuit: one on the main and the other on a subline with Jason Bourne's fingerprint. The bomb on the main line kills the power while Kirill kills the agent and the source, and steals the files and money, which he gives to Russian oil magnate Yuri Gretkov. Kirill then travels to Goa to kill Bourne, but Bourne flees with Marie. As Bourne and Marie are driving away, Kirill fires a sniper rifle at the car that kills Marie, and the car veers off the bridge they are on into a river. Kirill leaves, believing that he killed Bourne. Bourne manages to swim away undetected, leaving Marie dead in the river (after unsuccessful mouth-to-mouth resuscitation). He then burns all of Marie's passports and pictures except for one picture showing both of them in an embrace. He also clears the house they lived in, taking all of the notes that Bourne used to dictate his dreams. Bourne travels to Italy to learn why he is again being targeted.
After Landy's team finds the planted fingerprint and determines that it is linked to the CIA's secret Treadstone project, Landy flies to CIA headquarters in Langley, Virginia to find out more information. In Virginia, she learns that it is Bourne's fingerprint, but CIA chief Ward Abbott claims he does not know Bourne's whereabouts. Landy tells Abbott that the CIA agent who is believed to have stolen the $20 million was named in the stolen files. A Russian politician, Vladimir Neski, was going to identify the thief, but before he could do so, he was murdered by his wife in a Berlin hotel, who then committed suicide. Landy believes Bourne killed the CIA officer and source in Berlin, so Abbott and Landy set out to capture him.
When Bourne arrives in Naples, Italy, he is detained by an Italian Carabinieri officer and a CIA interrogator, but Bourne incapacitates the two men with his reflexes and without trying also and, after copying the agent's SIM card on his own cell phone, escapes. This allows Bourne to listen in on a call from Landy; he learns her name and phone number, and finds out that he is suspected in the recent killings in Berlin. He steals a car and heads to Munich, where he interrogates the last remaining Treadstone agent, Jarda, who tells him that the project was shut down following Conklin's death. They have a struggle in which Bourne emerges victorious by strangling Jarda. He creates a leak in the house's natural gas valve and puts a magazine into the toaster to create an explosion that slows down the police and allows Jason to escape. Landy and Abbott intercept and debrief Nicky Parsons, Treadstone's handler in Paris. Bourne arranges to meet Nicky, whom he interrogates. He learns that Abbott was the head of Treadstone, not Conklin. Upon hearing their conversation over Nicky's transmitter, Landy begins to believe that Bourne is being framed. When Abbott's young associate Danny Zorn tells Abbott that he thinks Bourne was framed, along with having evidence to prove he was, Abbott kills him to silence this information. Bourne goes back to the Brecker Hotel in Berlin and remembers more about the Neski mission: he was sent to kill Neski on Conklin's orders; and when Neski's wife unexpectedly showed up, he shot both of them and made it look like a murder-suicide. Bourne then learns that Abbott and Gretkov stole the money, and that Abbott had ordered Kirill to kill Bourne, Neski's murder, the murder of the agents by Kirill, and the planted fingerprint.
Out of respect for Marie's belief in non-violence, Bourne spares Abbott's life. After Zorn's body is discovered, Landy confronts Abbott, who commits suicide. Landy later receives the tape Bourne recorded which implicates Abbott in the conspiracy. Bourne travels to Moscow, where he is shot and wounded by Kirill. After an extended car chase, Bourne forces Kirill's car to crash into a concrete divider in a tunnel. Kirill is shown to be extensively injured and incapacitated, and presumably dies. (This is later confirmed in the next film, where Marie's brother asks if Bourne killed him.) Gretkov is arrested by Russian police after Landy provides them with the evidence she got from Bourne. Bourne goes to the apartment of Vladimir Neski's daughter so she won't have to live thinking her mother killed her father. He admits that he killed them both on a mission gone wrong, and apologizes. The final scene (which actually takes place during the events of the next film "The Bourne Ultimatum" in which it is shown) jumps to New York City, where Bourne contacts Landy to ask why the CIA is still looking for him. After Landy thanks Bourne for supplying the tape, which the CIA has used to settle matters concerning his frame-up, she tells Bourne his real name (David Webb) and birth date (April 15, 1971). Bourne declines Landy's offer to come back to the CIA offices in person, and he fades into a crowded street scene.
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